[3] The three major axes of the barycentric subdivisions are: Further indicators incorporated into the system are: Biotemperature is based on the growing season length and temperature.
Holdridge's system uses biotemperature first, rather than the temperate latitude bias of Merriam's life zones, and does not primarily consider elevation directly.
[8] Some research suggests that under the scenario of continually increasing greenhouse gas emissions, known as SSP5-8.5, the areas responsible for over half of the current crop and livestock output would experience very rapid shift in its Holdridge Life Zones.
This includes most of South Asia and the Middle East, as well as parts of sub-Saharan Africa and Central America: unlike the more developed areas facing the same shift, it is suggested they would struggle to adapt due to limited social resilience, and so crop and livestock in those places would leave what the authors have defined as a "safe climatic space".
In contrast, under the low-emissions SSP1-2.6 (a scenario compatible with the less ambitious Paris Agreement goals, 5% and 8% of crop and livestock production would leave that safe climatic space.