Rivalries began as early was 393/2 BCE, with Satyros and his co-regent and brother Seleukos attempting to besiege Theodosia, but failing in their pre-attempt.
After settling his affairs in order and solidifying his throne, Leukon, Satyros's eldest son and successor, besieged Theodosia once more.
[8] Five years after his original siege, Leukon had noticed that Tynnichus and his forces had been recalled to Heraclea Pontica after Clearchus made himself tyrant.
The annexation of Theodosia allowed the Spartocids to use the port for their export of grain to the city-state of Athens, which would prosper under its trade with the Bosporan Kingdom.
The only later hostilities between the Bosporans and the Heracleotes is around the reign of Leukon II and his brother Spartokos IV, some time in 235 BCE.