[1] The tibia was discovered in 1993 by archaeologist Mark Roberts and his team of the Institute of Archeology at University College London.

[5] The two teeth found show scratches, suggesting an eating technique in which food was cut with a tool whilst gripped between the jaws.

[6] Boxgrove in this period evolved from coastal mudflats beneath chalk cliffs to a mosaic of habitats as the sea retreated.

The 2022 study has found that the teeth fit in the morphological range of the Sima de los huesos fossils, which have been identified by aDNA as early Neanderthals.

The tibia falls outside the Sima de los Huesos range, and would fit the last common ancestor of modern humans and Neanderthals.