Following transfer from First ScotRail to Northern Rail in 2011, the first-class seating was refitted to the DTS(A) vehicles; but it was removed again in 2015 and replaced with a universally-accessible toilet.

Until opening of the Stansted Airport spur the units worked peak-hour services between Cambridge and Liverpool Street, but also found use on special trains wandering as far afield and York and Colchester.

[10] In December 2001, all were transferred to fellow National Express franchise ScotRail to replace the slam-door Class 305s on the North Berwick Line.

[10] Having been deemed surplus by One, in July 2005 all returned to Scotland to operate with First ScotRail on the North Berwick Line,[19][20] while also being used on peak services to Glasgow and Carstairs.

[24][25] Based at Neville Hill TMD, they operated services from Leeds to Doncaster, Bradford Forster Square, Skipton and Ilkley.

[26][27] In July 2020 all were leased to Greater Anglia to operate services out of London Liverpool Street on the Great Eastern Main Line.