[3][4] The genus is native to the Palearctic, Afrotropical, Indomalayan and Australasian realms.

[3] Adults enter through the fig ostiole, a narrow, bract-lined passage, then pollinate and attempt to oviposit on the flowers.

The adult offspring emerge from the gall and mate in the fig, before the winged female wasps disperse, carrying the flower pollen with them.

[5] Sycophaga sycomori oviposits inside the short-style flowers, thereby stimulating the growth of endosperm tissue and the enlargement and ripening of the syconium which holds the wasp-bearing drupelets, without pollination taking place.

[6] The parasitic species Apocrypta guineensis and Sycoscapter niger use long ovipositors to pierce the fig wall to infect the larvae during their development inside the flower galls,[7] and consequently reduce pollinator production.