[1] The character associations are: Captain Wings and the Black Condor; Dyna-Mite and Doll Man; Ghost Girl and the Phantom Lady; the Spirit of '76 and Uncle Sam; Thunder Fist and the Human Bomb; and Tommy Lightning and the Ray.
The Crusaders first appear in the World War II title The Invaders and capture the crew of a crashed German bomber.
The Crusaders are revealed to be guided by a cab driver and apparent British spy called "Alfie", who can cancel their powers courtesy of a technological belt the character wears.
Dyna-Mite, who has no memory of his former life, becomes suspicious and spies on Alfie, learning that he is a Nazi agent and is planning to use the heroes to assassinate the king.
The Crusaders, now powerless, disband, although Dyna-Mite—revealed to be Roger Aubrey, the close friend of Brian Falsworth, the brother of Invader Spitfire—remains trapped in a minute form.