The story continues where Days of Blood and Starlight leaves off, and concerns the invasion of Earth by the Seraph Empire and the alliance between the Misbegotten.

When Zuzana and Mick come bearing news of Jael's arrival on Earth, the new Chimaera leadership is able to negotiate an alliance with the Misbegotten.

Eliza Jones is a 24-year-old researcher in Washington, D.C., where she is working on her Ph.D. As a girl, she escaped a cult that worshipped her as a prophet and descendant of the angel Elazael.

Akiva is discovered by Stelian emissaries who, following his accidental disruption of the fabric of the universe while casting a spell, intend to kill him.

On Earth, Karou, Akiva, Zuzana, Mik, and a Chimaera soldier discover Eliza, speaking prophecies in Seraphim, on their way to the Vatican.

However, using the same spell Akiva used to synchronise the burning of Brimstone's portals, they are able to set an incendiary charge on Jael and force him to return to Eretz unarmed.