The tiny fruit bodies of the fungus resemble minute, white cups that grow scattered or in groups on the bark of ash trees.
The single species of Episphaeria was originally described under the name Cyphella fraxinicola by Miles Joseph Berkeley and Christopher Edmund Broome in an 1875 publication.
The online taxonomical database MycoBank lists it as part of the Strophariaceae, while Index Fungorum classifies it in the Inocybaceae.
[10] The minute fruit bodies of Episphaeria fraxinicola are cyphelloid, meaning they resemble species of discomycetes (or "cup fungi") in the Ascomycota.
The fruit bodies consist of caps that are 0.25–2 mm, white, circular or nearly so, and lay flat on the substrate without a stem.