Europelta

[1] Europelta has an estimated length of 5 metres (16 feet) and weight of 1.3 tonnes (2,866 lbs), making it the largest member of the clade Struthiosaurini.

[2] In 2011, the remains of two partial skeletons of an ankylosaur were discovered from the Fundación Conjunto Paleontológico of Teruel-Dinópolis locality AR-1 of the lower, located east of Ariño, in the northern Teruel Province in the Community of Aragón.

The holotype specimen, AR-1/10, consists of a mostly complete skull, isolated nasals, a dentary fragment, isolated teeth, an atlas, cervical vertebrae, cervical ribs, dorsal vertebrae, a section of synsacrum, isolated dorsal ribs, dorsal rib fragments, caudal vertebrae, chevrons, a coracoid with a small portion of scapula, a scapular blade fragment, xiphosternal plates, partial humeri, an ischium, an ilium, pubes, and 70 osteoderms.

[1] In 2016, Gregory S. Paul gave Europelta an estimated length of 5 metres (16 feet) and a weight of 1.3 tonnes (2,866 lbs).

[2] Kirkland and colleagues (2013) diagnosed Europelta based on the quadrate being shorter and mediolaterally wider than in any other ankylosaur, the concave shape of the posterior margin of the skull in dorsal view, the sacrum in lateral view is arched dorsally by 55°, the pubis is fused to the ischium and has a slot-shaped foramen between the post-pubic process and the orientation of the pubic peduncle which forms an ischiopubis, the tibia is longer relative to the length of the femur than in other ankylosaurs and the presence of laterally compressed, flanged osteoderms with a flat plate-like base that are present towards the front of the pelvic shield.

The palate does not have a pronounced hourglass appearance as in derived nodosaurids due to medially deflected tooth rows.

The lateral wall of the skull extends behind the orbit unlike those of polacanthids and most nodosaurids except for Peloroplites, Silvisaurus and Struthiosaurus.

The sheet-like pterygoids flexed nearly dorsally against the front portion of the basicranium as in nodosaurids and not open transversely as in ankylosaurids or polacanthines.

The skull roof is roughened texturally by remodelling of the bone surface and preserves skin impressions on the margins that are represented by shallow grooves.

The lateral margin of the angular is highly rugose, due to the texture and remodelling required to support a large scale.

The anterior centra has ventral sides that are characterized by two oriented paired fossae that extend from the front to back and are separated by a low keel.

The dorsal vertebrae near the front have large cylindrical centra that are flat at both ends which lack a constricted abdominal keel with circular neural canals and fused ribs.

The chevrons situated nearer to the centre are approximately as long as the neural spines but lack fusion to their respective caudal vertebrae.

The most distal four caudal vertebrae and their chevrons are fused together which form a tapering, terminal rod of bone at the end of the tail.

The femoral head has an articular surface directed dorsally and forms an angle of about 115° with the long axis of the femur.

Osteoderms with sharp anterior and posterior edges, and are hollow, asymmetric-based plate-like ran down the sides of the tail but decreased in size more rapidly.

Asymmetric, diamond to tear-drop shaped osteoderms are also found in both specimens and would have been in oblique rows anterior to the sacrum based on close relatives.

Medium to large oval to circular osteoderms were probably present on the posterior portion of the sacrum or within more expansive spaces among the larger dorsal armour.

The posterior margin of the ilium preserves large, subequal-sized osteoderms that are tightly sutured together but was not coossified as in Aletopelta.

[1] Kirkland et al. (2013) originally classified Europelta as a member of Struthiosaurinae but did not conduct a phylogenetic analysis to determine its exact relationships within the clade.

They suggested that this was not a coincidence but an indication that the fauna of North America and Europe had not yet been separated by the developing northern Atlantic Ocean, as had been presumed earlier.

[6][7] Sauroplites Mymoorapelta Dongyangopelta Gastonia Gargoyleosaurus Hoplitosaurus Polacanthus Peloroplites Taohelong Sauropelta Acantholipan Nodosaurus Niobrarasaurus Ahshislepelta Tatankacephalus Silvisaurus

[8] Due to the absence of arenaceous foraminifera and dinoflagellates, the member was formed well inland of marine and brackish water influences.

The layer that the holotype and paratype specimens of Europelta were obtained from have a characteristic dark olive-gray to olive-black mudstone that preserves a high percentage of fossil plant debris.

[1] The Escucha Formation was inhabited by various species of ostheicthyians, freshwater bivalves, gastropods, arthropods, turtles, and crocodylomorphs such as Hulkepholis, Anteophthalmosuchus and an indeterminate goniopholidid.

Similar assemblages of charcoalified ferns have been recorded in several localities from England that represent fluvial to lacustrine deposits of Hauterivian to Barremian age.

This taphonomic plant assemblage indicates the action of a local fire over the vegetation growing in a nearby fluvial plain.

The combination of the properties of the burnt soil and the action of heavy rainfalls suggests the charred remains moved downstream and were buried later in a near depositional area.

Size comparison of Europelta and a human.
Mandible of the paratype specimen.
Cervical vertebrae and ribs of holotype.
Sacrum of the paratype.
Femur, tibia, fibula and calcaneum of the paratype.
Larger osteoderms of holotype.
Dorsal osteoderms of holotype.
Skeletal reconstruction of Europelta .
Paleogeographic reconstruction of the Escucha outcrop belt.