Formiciinae is an extinct subfamily of ants known from Eocene deposits in Europe and North America.

Titanomyrma also contains a third species, T. lubei described in the same paper as the genus and which is the second member of the subfamily known from North America.

Only the queens of Dorylus wilverthi currently reach similar lengths, up to 52 millimetres (2.0 in) as the smallest species of Titanomyrma, T.

[1] Formiciinae members were restricted in habitat to living in regions which had a mesic wet climate and an average mean annual temperature of 20 °C (68 °F) or higher.

While the average temperatures for this route are thought to have been lower than the range needed for Formiciinae species, a series of warmer events throughout the Eocene are suggested as aides in the crossing.