Sphecomyrminae

See text Sphecomyrminae is an extinct subfamily of ants in family Formicidae known from a series of Cretaceous fossils found in North America, Europe, and Asia.

The tribe Sphecomyrmini contains the six genera Armania, Cretomyrma, Gerontoformica, Orapia, Pseudarmania and Sphecomyrma; while Zigrasimeciini contains Boltonimecia and Zigrasimecia.

Sphecomyrminae is the most basal of the Formicidae subfamilies, but has not been included in several phylogenetic studies of the family.

The subfamily is characterized by three major synapomorphies, the short pedicel, a second flagellar segment that is double the length of the other antenna segments, and the loss of the apical end of the CuA veins in the wings of adult males.

[1] Further review of the haidomyrmecins resulted in them being elevated to the separate subfamily Haidomyrmecinae and removed from Sphecomyrminae.