He wrote among professors such as Lothar Geitler and Tschermak Woess about the "evolution and karyology of elaiosomes to fruit and seeds."
In 1972, he received a Ph.D. From 1970, he was a research associate at the Upper Austrian Provincial Museum Linz, initially as Head of botany and invertebrates, from 1985 as director deputy of the National Museum and 1990 to 1991 as interim director.
From 1993 to 2003 he was head of the newly founded Biology Centre of the National Museum.
In 1982, he received the venia legendi of Systematic Botany at the University of Salzburg.
Other subjects are the tribes Antirrhineae the Scrophulariaceae, and Pinguicula (Lentibulariaceae).