Galulatherium is an extinct genus of possibly gondwanathere mammal, from the Late Cretaceous (Turonian-Campanian)-aged Galula Formation of Tanzania.

First described in 2003, TNM 02067 has been tentatively identified as a sudamericid—an extinct family of high-crowned gondwanathere mammals otherwise known from South America, Madagascar, India, and Antarctica.

The describers could not exclude other possibilities, such as that the jaw represents some mammalian group known only from younger, Cenozoic times (less than 66 million years ago).

[1] Galulatherium was discovered in 2002[2] in the locality TZ-07 in the Mbeya Region of southwestern Tanzania, which has also yielded remains of various other vertebrates, including birds and other saurischian dinosaurs.

Age estimates for the RSU have ranged from middle Jurassic to Miocene, but according to Krause and colleagues, part of this discrepancy is the result of confusion between two superficially similar rock units that outcrop nearby; the older one, where TZ-07 is located, is undoubtedly Mesozoic and the younger is Cenozoic.

Based on the presence of non-avian dinosaurs and osteoglossomorph fishes, Krause and colleagues assigned TZ-07 to the Cretaceous (146–66 million years ago).

[4] In 2007, Nancy Stevens and colleagues identified the unit that produced TNM 02067 as likely belonging to the middle part of the Cretaceous (around Aptian to Cenomanian).

[7] All the teeth are incomplete or absent, and lack both enamel and cementum, but what remains indicates that there was a large incisor at the front and five cheekteeth further back, separated by a diastema (gap) of about 2.5 mm (0.098 in).

[8] The dentary superficially resembles that of various other mammalian groups with enlarged incisors, such as rodents, lagomorphs, hyraxes, wombats, the aye-aye, and the extinct apatemyids, tillodonts, and taeniodonts—all of which are known only from the Cenozoic, less than 66 million years ago.

Sudamerica has only four, not five, cheekteeth (all of which are molariform), a higher, narrower incisor with a root that extends further through the dentary, and a shorter diastema; in all these respects, TNM 02067 is more primitive.