[1] The salt glands of mangroves such as Acanthus, Aegiceras, Aegialitis and Avicennia are a distinctive multicellular trichome, a glandular hair found on the upper leaf surface and much more densely in the abaxial indumentum.
Development of the glands resembles that of the nonglandular hairs until the three-celled stage, when the short middle stalk cell appears.
These cavities come to contain large amounts of cannabidiol in hemp-producing strains, or tetrahydrocannabinol and cannabinol in other cannabis plants.
These produce spheres of oily secretions, including terpenes, which pass through the cell membrane and wall to accumulate as vesicles in the secretory cavity.
The glands gradually darken as they mature, with loss of cannabinoids over time (perhaps to evaporation) and eventually undergo abscission from the plant.