[1] She began her PhD at the same institution later that year, based at the university’s Department of Animal Breeding, then under the directorship of noted pioneering geneticist Francis Albert Eley Crew.
She was awarded her doctorate in 1925 for her thesis researching the individual wool types and yield produced by the offspring of the Department’s ovine hybridisation programme.
[1] In 1928, Blyth was a keynote speaker at the 5th meeting of the Royal Physical Society of Edinburgh chaired by zoologist Marion Newbigin, where she delivered an illustrated paper on her early research endeavours relating the gonadic structure of fowl to their comb-growth.
[4] In July 1930, Blyth and Greenwood presented to the World's Poultry Congress in London a joint paper on their ongoing work at the University of Edinburgh Animal Breeding Research Department titled “Some Experiments Relating to the Ovarian Function in the Fowl”.
[5] When Crew departed from Edinburgh on active service during World War Two, Greenwood became acting director of the University’s Institute of Animal Genetics, including its Poultry Section.