He currently serves on the board of directors of the Open Source Applications Foundation and Code for America.

[3] Lilly co-founded[5] and served the CTO for Reactivity,[6] a start-up focused on consulting on technically difficult websites and incubating new companies.

Lilly, previously Mozilla's chief operating officer, succeeded Mitchell Baker as CEO in January 2008.

[10] Lilly left Mozilla's board of directors in March 2014, reportedly over the appointment of Brendan Eich as CEO.

His investments include Instagram,[12] Dropbox,[12] Tumblr,[12] Quip,[13] and MessageMe.