John Thoday

He was Arthur Balfour Professor of Genetics at Cambridge University between 1959 and 1983 and was elected a Fellow of the Royal Society in 1965.

During World War II he served in the RAF as a photographic intelligence officer.

He has published an important thesis on the meaning of biological progress in evolution and the role of genetic variation in determining long term fitness.

He has pioneered a method for the location on chromosomes of genes mediating continuous variation, and showed (contrary to accepted theory) that the genes at different loci affected the quantitative character in qualitatively different ways.

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