He served as a professor of hydrology at St. Petersburg University and took part in a number of oceanographic expeditions.
A year before his graduation, he took part in an expedition to the White Sea which made him interested in marine life and oceanography.
He became a lecturer at St Petersburg University in 1917 and became a professor in 1919 while also working at the State Hydrological Institute.
He took an interest in marine life and worked on a number of taxa, collecting and examining their distributions.
He identified faunal barriers formed by warm shallow waters in the Gorlo Strait.