His discoveries became a crucial point in negotiations over pacific salmon fisheries, as it could now be determined where each individual fish spawned, in the rivers of Canada or the United States.

He joined the Pacific Biological Station in Nanaimo, British Columbia, where was a government scientist, advisor, and diplomatic representative.

He became Head of the Fish Health and Parasitology Section of the Station in 1967 and was appointed Senior Scientist in 1990.

He suffered a heart attack in 1997 while walking home from work and died several days later, at the age of 69, after being airlifted to a Vancouver hospital.

The paper[2] contains the definition of widely used terms such as prevalence and has subsequently been cited thousands of times.