Phylogeny of pterosaurs

Preondactylus Dimorphodontidae Anurognathidae Campylognathoididae Rhamphorhynchinae Scaphognathinae Istiodactylus Ornithocheiridae Nyctosaurus Pteranodontinae Gallodactylidae Pterodactylus Lonchodectes Ctenochasmatidae Germanodactylus Dsungaripteridae Tapejara Tupuxuara Azhdarchidae The matrix includes 39 valid pterosaur species, although Rhamphorhynchus longicaudus and Nyctosaurus bonneri are usually considered to be synonymous with R. muensteri and N. gracilis respectively, plus a three outgroup species (Ornithosuchus longidens, Herrerasaurus ischigualastensis and Scleromochlus taylori).

Anurognathus ammoni Batrachognathus volans Dendrorhynchoides curvidentatus Sordes pilosus Scaphognathus crassirostris Preondactylus buffarinii Dorygnathus banthensis Dimorphodon macronyx Peteinosaurus zambellii "Eudimorphodon" rosenfeldi Campylognathoides liasicus Eudimorphodon ranzii Rhamphorhynchus longicaudus Rhamphorhynchus muensteri Pterodactylus antiquus Pterodactylus kochi Germanodactylus cristatus Germanodactylus rhamphastinus Ctenochasma Pterodaustro guinazui Cycnorhamphus suevicus Gallodactylus canjuersensis Nyctosaurus bonneri Nyctosaurus gracilis Pteranodon longiceps Istiodactylus latidens Coloborhynchus clavirostris Tropeognathus mesembrinus Anhanguera blittersdorffi Anhanguera piscator Anhanguera santanae Dsungaripterus weii Noripterus complicidens "Phobetor" parvus Tupuxuara leonardii Tapejara wellnhoferi Tapejara imperator Azhdarcho lancicollis Quetzalcoatlus sp In 2010, Brian Blake Andres wrote a review of pterosaur phylogeny in his dissertation.

[2] This phylogenetic analysis was used by Richard J. Butler, Stephen L. Brusatte, Brian B. Andres and Roger B. J. Benson (2012) to assess the morphological diversity and fossil sampling biases of the Pterosauria.

[4] An updated and more resolved version of this phylogeny was published formally by Andres and Timothy Myers in 2013, containing 185 characters and 109 ingroup taxa.

[5] Preondactylus buffarinii Austriadactylus cristatus Peteinosaurus zambellii Caviramus schesaplanensis Raeticodactylus filisurensis Eudimorphodon cromptonellus Eudimorphodon ranzii Eudimorphodon rosenfeldi Dimorphodon macronyx Parapsicephalus purdoni Campylognathoides liasicus Campylognathoides zitteli Scaphognathus crassirostris Dorygnathus banthensis Cacibupteryx caribensis Nesodactylus hesperius Rhamphorhynchus muensteri Harpactognathus gentryii Angustinaripterus longicephalus Sericipterus wucaiwanensis Sordes pilosus Pterorhynchus wellnhoferi Darwinopterus modularis Wukongopterus lii Changchengopterus pani Dendrorhynchoides curvidentatus Jeholopterus ningchengensis Anurognathus ammoni Batrachognathus volans Normannognathus wellnhoferi Germanodactylus cristatus Germanodactylus rhamphastinus Pterodactylus antiquus Pterodactylus kochi Ardeadactylus longicollum Boreopterus cuiae Feilongus youngi Cycnorhamphus suevicus Gallodactylus canjuersensis Kepodactylus insperatus Elanodactylus prolatus Huanhepterus quingyangensis Plataleorhynchus streptophorodon Gnathosaurus subulatus Gnathosaurus macrurus Ctenochasma elegans Ctenochasma roemeri Pterodaustro guinazui Eosipterus yangi Beipiaopterus chenianus Gegepterus changi Haopterus gracilis Muzquizopteryx coahuilensis "Nyctosaurus" lamegoi Nyctosaurus gracilis Alamodactylus byrdi Pteranodon longiceps Pteranodon sternbergi Longchengpterus zhaoi Nurhachius ignaciobritoi Liaoxipterus brachyognathus Istiodactylus latidens Istiodactylus sinensis Lonchodectes compressirostris Aetodactylus halli Cearadactylus atrox Brasileodactylus araripensis Ludodactylus sibbicki Liaoningopterus gui Anhanguera araripensis Anhanguera blittersdorffi Anhanguera piscator Anhanguera santanae Tropeognathus mesembrinus Ornithocheirus simus Coloborhynchus clavirostris Coloborhynchus wadleighi Bennettazhia oregonensis Nemicolopterus crypticus "Sinopterus" gui Huaxiapterus jii Eopteranodon lii Sinopterus dongi "Huaxiapterus" corollatus "Huaxiapterus" benxiensis Bakonydraco galaczi Tapejara wellnhoferi Tupandactylus navigans Tupandactylus imperator Eoazhdarcho liaoxiensis Shenzhoupterus chaoyangensis Chaoyangopterus zhangi Jidapterus edentus Radiodactylus langstoni Azhdarcho lancicollis