See text The Lycopodiaceae (class Lycopodiopsida, order Lycopodiales) are an old family of vascular plants, including all of the core clubmosses and firmosses, comprising 17 accepted genera[2] and about 500 known species.
The spores are of one size (i.e. the plants are isosporous) and are borne on a specialized structure at the apex of a shoot called a strobilus (plural: strobili), which resembles a tiny battle club, from which the common name derives.
In 2016, Field et al. proposed that the primary division is between Lycopodielloideae plus Lycopodioideae and the Huperzioideae (names sensu PPG I).
Field et al. (2016) say "Most Lycopodiaceae species have been re-classified into different genera several times, leading to uncertainty about their most appropriate generic identification.
"[9] In the PPG I system, the family has 16 accepted genera, grouped into three subfamilies, Lycopodielloideae, Lycopodioideae and Huperzioideae, based in part on molecular phylogenetic studies.
The Huperzioideae differ in producing spores in small lateral structures in the leaf axils,[9] and it has been suggested that they be recognized as a separate family.
The two subfamilies Lycopodioideae and Huperzioideae diverged ~350 million years ago, but has evolved so slowly that about 30% of their genes are still in syntenic blocks (remaining in the same arrangement).