He is a member of the Department of Molecular Biology and of the Lewis-Sigler Institute for Integrative Genomics, where he is currently director of graduate studies.

Working with Yigal Meir, Wingreen formulated the Meir-Wingreen Formula which describes the electric current through an arbitrary mesoscopic system.

[2] Wingreen then received his Ph.D. in theoretical condensed matter physics from Cornell University in 1989 as a Hertz Fellow.

There, along with Yigal Meir, he formulated the Meir-Wingreen Formula that describes the electric current through an arbitrary mesoscopic system.

[5] Wingreen's current research focuses on modelling intracellular networks in bacteria and other microorganisms, as well as studies of microbial communities.