[1] Ecosystems employ biodiversity in the food webs that recycle natural materials, such as mineral nutrients, which includes water.
From a practical point, it does not make sense to assess a terrestrial ecosystem by considering the full column of air above it as well as the great depths of Earth below it.
While an ecosystem often has no clear boundary, as a working model it is practical to consider the functional community where the bulk of matter and energy transfer occurs.
[11] Ecosystems execute closed loop recycling where demand for the nutrients that adds to the growth of biomass exceeds supply within that system.
There are regional and spatial differences in the rates of growth and exchange of materials, where some ecosystems may be in nutrient debt (sinks) where others will have extra supply (sources).
[6][12] In terms of a food web, a cycle or loop is defined as "a directed sequence of one or more links starting from, and ending at, the same species.
"[17] Different ecosystems can vary in their recycling rates of litter, which creates a complex feedback on factors such as the competitive dominance of certain plant species.
Different rates and patterns of ecological recycling leaves a legacy of environmental effects with implications for the future evolution of ecosystems.
Because the earthly pool of these elements is limited and the rates of exchange among the various components of the biota are extremely fast with respect to geological time, it is quite evident that much of the same material is being incorporated again and again into different biological forms.
Organic farms that employ ecosystem recycling to a greater extent support more species (increased levels of biodiversity) and have a different food web structure.
The persistent legacy of environmental feedback that is left behind by or as an extension of the ecological actions of organisms is known as niche construction or ecosystem engineering.
'"[39] The basic idea of a balance of nature, however, can be traced back to the Greeks: Democritus, Epicurus, and their Roman disciple Lucretius.
Dumas and Boussingault (1844) provided a key paper that is recognized by some to be the true beginning of biogeochemistry, where they talked about the cycle of organic life in great detail.
[53][54] This idea was similarly expressed in 1954 by ecologist Paul Sears: "We do not know whether to cherish the forest as a source of essential raw materials and other benefits or to remove it for the space it occupies.
[56] Rachel Carson was an ecological pioneer in this area as her book Silent Spring inspired research into biomagnification and brought to the world's attention the unseen pollutants moving into the food chains of the planet.
[62]: 62 Microplastics and nanosilver materials flowing and cycling through ecosystems from pollution and discarded technology are among a growing list of emerging ecological concerns.
[65] Human wastes in the Anthropocene are creating new systems of ecological recycling, novel ecosystems that have to contend with the mercury cycle and other synthetic materials that are streaming into the biodegradation chain.
[66] Microorganisms have a significant role in the removal of synthetic organic compounds from the environment empowered by recycling mechanisms that have complex biodegradation pathways.
The effect of synthetic materials, such as nanoparticles and microplastics, on ecological recycling systems is listed as one of the major concerns for ecosystems in this century.