Overoraptor (/ˌʊuvərʊuˈræptər/, meaning "piebald thief") is an extinct genus of paravian theropod of uncertain affinities from the Late Cretaceous Huincul Formation of Argentinian Patagonia.
The fossil specimens of Overoraptor were unearthed by Dr. Roberto Nicolás Chimento in the Violante Farm locality, which is southeast of the Ezequiel Ramos-Mexía lake, in the northwestern portion of Rio Negro Province, Argentina.
[2] Motta and colleagues diagnosed Overoraptor as being distinct from all other paravians from the following unique autapomorphies: a medial deflection at the distal end of the scapula, a reduced and ridge-like acromial process, an extensive crest on metacarpal I, a crest ending in a distal posterior tubicle on metatarsal II, and a metatarsal III that is taller than it is wide at the midpoint along its length.
[1] The scapular anatomy of Overoraptor resembles that of stem-avialans in the presence of a cup-shaped glenoid fossa, but it differs from most other paravians in that the acromial process projects dorsally rather than medially.
The overall morphology of the scapula resembles the enigmatic paravian taxon Rahonavis because the scapular blade projects medially outward at its distal-most end.
[1] The metacarpal bones of Overoraptor were noted by Motta and colleagues to be unusual in shape, namely that it is proportionally much wider and shorter than those known from other non-avialan paravians and there is a distinctly asymmetrical ginglymoidal surface which articulates with the lateral condyle.
The Overoraptor-Rahonavis clade was recovered with the following synapomorphies: a complex set of ridges on the lateral surfaces of the caudal centra and a medial deflection of the distal end of the scapular blade.
Even considering the similarities between the two taxa, Motta and colleagues also remark that the recovery of this clade may be the result of the unique mix of plesiomorphic and derived characters present in Overoraptor.
[1] Dromaeosauridae Microraptoria Unenlagiidae Overoraptor Rahonavis Alcmonavis Jeholornithidae In 2021, Andrea Cau and Daniel Madzia published their redescription of the troodontid Borogovia, which contained a phylogenetic analysis using a wide variety of coelurosaur taxa.
They used several different interpretive frameworks including the coding of Rahonavis as an unenlagiine, Balaur as a velociraptorine, and halszkaraptorines as being basal to the troodontid-dromaeosaurid split as well as an unweighted analysis without these constraints.
A study of the El Zampal section of the formation found hornworts, liverworts, ferns, conifers, and some angiosperms (flowering plants).
[18] The Huincul Formation is among the richest Patagonian vertebrate associations, preserving fish including lungfish and gar, chelid turtles, squamates, sphenodonts, neosuchian crocodilians, and a wide variety of dinosaurs.
[20] The Violante Farm locality is one of the most productive in the Huincul Formation, having yielded specimens of a wide variety of non-avian dinosaur species.