[4] The genus was originally defined based on characteristics of adult males: deflexed elytral apex, trisinuate ventrite 7, and a metafemoral comb.

Developments in molecular sequencing techniques have improved the ability of entomologists to identify and delineate new species based on only female or larval specimens that were not collected in association with males.

The males of two species, P. malaccae and P. tener, are often found in large groups on certain trees and perform synchronized flashing displays to attract females for mating.

[1] Many of the species examined by Jusoh et al. in 2018 using genetic sequencing of the mitochondrial DNA were associated with mangroves, as their larva need a continuously damp environment to develop.

This form of ecotourism has been used to stress the importance of mangrove protection and preservation, as many of the tour operators are local fishermen earning supplemental income.