Punatitan

Punatitan (meaning "puna giant") is a genus of titanosaurian sauropod dinosaur from the Late Cretaceous Ciénaga del Río Huaco Formation of La Rioja, Argentina.

The specific name refers to geologist Tim Coughlin, who first reported dinosaur fossils in the discovery locality.

Known from the holotype CRILAR-Pv 614 (Paleovertebrate Collection of Centro Regional de Investigaciones Científicas y Transferencia Tecnológica de La Rioja, Argentina), a partial skeleton composed of the anterior portion of posterior cervical vertebra (likely C12), two middle dorsal vertebrae (likely D6–D7), a partial sacrum, 13 articulated caudal vertebrae (some with articulated haemal arches), the right pubis, the left ischium, and several dorsal ribs, Punatitan was roughly 14 metres (46 ft) long.

[1] The describers' phylogenetic analysis places Punatitan as a derived member of the Lithostrotia, in the clade Aeolosaurini, which they recover as a subclade of Rinconsauria, opposite to other analyses.

Their cladogram is shown below: Malawisaurus Epachthosaurus Dreadnoughtus Rapetosaurus Isisaurus Tapuiasaurus Saltasauridae Baurutitan Bonitasaura Notocolossus Lognkosauria Rinconsaurus Muyelensaurus Overosaurus Arrudatitan Punatitan Aeolosaurus Bravasaurus Trigonosaurus Gondwanatitan Uberabatitan The holotype locality, the Quebrada de Santo Domingo site, preserves one of the largest concentrations of titanosaur eggs in the world.