(doctorate) degree from the Oxford University, where he had mentors like Nobel laureates Sir Hans Krebs and Rodney Porter.
[3] He has been fellow of the Wellcome Trust (1972–1974) and the Biochemical Society, UK (1972–1997), other than holding numerous other honors, awards, and fellowships.
[5] In past, he had served as a Distinguished National Professor, Higher Education Commission Pakistan, and director general of the Biomedical & Genetic Engineering Division, Islamabad,[2] at the Institute for Biotechnology & Genetic Engineering, University of Karachi, Pakistan.
of Chemistry, Karachi University, 1993-, The World Bank Committee for Higher Education in Pakistan, elected member, Scientific and Medical Advisory Board, the International Retinitis Pigmentosa Society (Retinitis Pigmentosa International) and The Foundation Fighting Blindness society, Washington DC (1993–present), Human Genome Organization (HUGO), London (UK), founding member, Human Genome Diversity Project (HGDP) USA.
He has written over two hundred publications, including papers in Science, Nature Genetics and Genome Research.