Richard M. Harland FRS[3][4] is CH Li Distinguished Professor of Genetics, Genomics and Development at the University of California, Berkeley.

Harland completed his PhD at the Medical Research Council (MRC) Laboratory of Molecular Biology (LMB) at the University of Cambridge under the supervision of Ron Laskey on regulation of DNA replication in Xenopus embryos.

[1][5] Following postdoctoral work at the LMB and at the Fred Hutchinson Cancer Research Center with Harold M. Weintraub and Steve McKnight in Seattle he moved to the University of California, Berkeley in 1985.

Major contributions include understanding the early embryo, and the induction and patterning of the neural plate and the Spemann-Mangold organizer.

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