[1] He published more than 130 research papers[2] that helped advance the study of plant–insect interactions, interspecific competition, predator prey interactions and food web dynamics.

He studied the ecology of sap-feeding insects, both in natural and cultivated settings.

His personal worldwide collection of butterflies numbered over 36,000 specimens.

[1] His butterfly collection was donated, at least in part, to the McGuire Center for Lepidoptera and Biodiversity.

[3] While at College Park, one of his doctoral students was Lawrence Hanks.