[13] Williams’s plan was to continue working with Irving for his DPhil, but he first visited the lab of Arne Tiselius at Uppsala University.

Come back again in two weeks to give me your decision.”[14] Williams joined Wadham College, Oxford in 1955 and remained there for the rest of his life.

[19] In the first paper ever published in the Journal of Theoretical Biology Williams argued that spatial separation of the H+ and OH– ions produced in the conversion of ATP to ADP would be necessary for the catalysis.

Williams was appointed Member of the Order of the British Empire (MBE) in the 2010 New Year Honours for services to the community in North Oxford.

[27] He was elected a Fellow of the Royal Society (FRS) in 1972[14] and was a Foreign Member of the Swedish, Portuguese, Czechoslovakian and Belgian science academies.

Jelly read English language and literature at St Hilda's College, Oxford between 1952 and 1955, but the birth of their first son, Timothy Ivor, interrupted her final exams.