He is the leader of the Lupines, a species resembling humans that he claims evolved from canines instead of primates through convergent evolution.

Created by writer Jeph Loeb and artist Simone Bianchi, the character first appeared obscured by shadow in Wolverine (vol.

He is shown leading the Lupines in prehistoric[2] and barbaric[3] times, being an emperor in Ancient Rome,[4] as well as being the main force behind Weapon X.

Nick Fury revealed to Wolverine that Romulus has been manipulating people from his family's bloodline for centuries, and that he plans for Daken to become the next-generation Weapon X.

Romulus told Daken to kill all the Weapon X experiments that gave them a healing factor with his Muramasa claws.

[9] After Wolverine temporarily subdues Omega Red, he is captured by Wild Child, who reveals that Romulus made Logan and several villains battle each other, his agenda being vaguely described as him needing a successor.

[volume & issue needed] After his confrontation with Wolverine he coordinates a successful assassination in Tokyo from what seems to be his base of operations, a European-style castle.

[volume & issue needed] Wolverine decides to confront Romulus one last time, using the help of various other heroes and his son Daken.

In the end, as Daken is about to kill Romulus, Logan has Cloak teleport him into the Dark Force dimension.

For a second Wolverine thinks he should behead Romulus, just like he did with Creed, but in the end he decides to leave him stranded in the Dark Force dimension.

After a losing fight against Romulus, a mysterious woman with long red hair appears and tells Wolverine that the answers that he seeks are at the Weapon X facility.

She further reveals that the whole "Lupine Sapiens" story was all just an elaborate hoax by her brother intended to be a ruse for his real goal of creating a master race of natural mutants artificially enhanced by a new type of adamantium created using vibranium illegally mined from Wakanda, using Wolverine as a template.

Like Wolverine, Romulus' primary power is an accelerated healing factor that allows him to rapidly regenerate from injury.

Romulus' healing factor grants him virtual immunity to known diseases and renders him highly resistant to most forms of drugs.

It also affords him greatly extended longevity by slowing his natural aging process to an extraordinary degree as he claims to be tens of thousands of years old.

His healing factor grants him partial immunity to the fatigue toxins generated by his muscles during physical activity, so he has much greater stamina and endurance than an ordinary human.

Although he often appears naked, he has been shown to wear clothes on several occasions, possibly making use of the fact that only a few living people can recognize him.

He also has extensive knowledge of many foreign and ancient cultures as he has alleged to have participated in the rise and fall of many civilizations throughout his existence.