Santanadactylus

UvA M 4895, consisting of two cervical vertebrae from a different individual, was assigned as a paratype, referred because it was found in the same lot of 25 chalk nodules bought from collectors.

[3][4] In 1985, Peter Wellnhofer, a German paleontologist who has written numerous scientific publications on pterosaurs, named three additional species: S. araripensis, S. pricei, and S.

S. pricei, named after Llewellyn Ivor Price, was the smallest of the three species; it was based on BSP 1980 I 122, a left wing from the elbow down, and additional arm material has been referred to it over the years.

[4] S. spixi, intermediate in size, was based on BSP 1980 I 121, another left wing, the name honoring Johann Baptist von Spix.

[8] Averianov (2014) considered Santanadactylus spixi a nomen dubium probably synonymous with Tupuxuara longicristatus, and he also indicated that the paratype of S. brasiliensis was likely referable to T.

Wellnhofer (1991) considered it a member of the Ornithocheiridae based on the structure of the humerus, but noted that the elongation of the paratype neck vertebrae distinguished it from other ornithocheirids.

Reconstructed skeleton, Wyoming Dinosaur Center