Smilodectes is a genus of adapiform primate that lived in North America during the middle Eocene.

Smilodectes has a small cranium size and the foramen magnum was located at the back of the skull, on the occipital bone.

[1] Smilodectes gracilis was an adapiformes primate from the early Eocene, some 55 million years ago.

[2] S. gracilis had a dental formula of 2.1.4.32.1.4.3 and had a relatively short snout, with rounded frontal bone as compared to other nothactines.

[2] Based upon its postcranial skeleton, S. gracilis was a vertical clinger and leaper.