The manga's plot focuses on Sunako Nakahara, a girl who was called ugly by the first boy she had a crush on and isolated herself from life, losing her sociability and care for her physical appearance.
Four handsome boys, Kyouhei, Takenaga, Yuki, and Ranmaru must try to turn her into a lady to please her aunt, who is also their landlady, with the promise that they won't have to pay the rent if they succeed.
She hates excessive light exposure and does poorly in hot weather, though the latter is because she grew up in northern Japan.
She has three "friends"; a skeleton called Josephine (with her husband, sergeant George) and two anatomical figures named Hiroshi-Kun and Akira-Kun.
While she suffers from severely low self-esteem, she's quite beautiful (most of her physical imperfections are a result of her self-imposed isolation from the world) and is a frighteningly tough fighter, being able to lift someone twice her size with one hand.
She is portrayed (mostly) throughout the manga in chibi form as a sort of visual metaphor for how she and others see herself; only becoming full-sized and in proportion with the others when her mood becomes bold or confident.
Recent chapters have also shown her forgetting her first crush and becoming more open to others, including her peers and teachers at school.
Voiced by: Showtaro Morikubo (Japanese); Jessie James Grelle (English) Live-Action: Kazuya Kamenashi The male protagonist of the series.
While he says that all he cares about is getting free (or the lowest amount required to be paid) rent, he has a soft spot for Sunako and empathizes with her more so than the others.
He is the most popular of the boys, and always has trouble keeping down a part-time job because his fans will chase after him, or his boss (male or female) will sexually harass him.
Since Sunako is the only girl who can understand his true personality regardless of his good looks, he gradually develops an obvious attraction towards her, despite being dense towards his feelings.
Voiced by: Tomokazu Sugita (Japanese); Vic Mignogna (English) Live-Action: Hiroki Uchi The most intellectual of the four, Takenaga is called the "Boss" (a pun on his name in Japanese, which can be read as buchou, meaning "boss") by his friends and is the least fazed by Sunako's morbid habits.
Voiced by: Yamauchi Yuuya (Japanese); Greg Ayres (English) Live-Action: Yuya Tegoshi The most androgynous of the four, "Yuki" is gentle and is initially the most frightened of Sunako.
He is usually responsible for choosing what sort of clothes Sunako wears when she is forced to dress up and is considered to have a good sense of fashion.
Voiced by: Hirofumi Nojima (Japanese); Chris Patton (English) Live-Action: Shuntaro Miyao A ladies' man, Ranmaru is often going out with various beautiful older (and often married) women.
The very sight of Sunako initially has him breaking out in hives, but when her true beauty is revealed, he decides that she has "potential" to become a real lady.
Like Takenaga, he comes from a wealthy family (who own a series of hotel chains) but was sent to live with the Landlady because he exploited his wealth and his parents wanted him to become normal.
In hopes of changing his womanizing habits, his parents engage him with a reserved and conservative girl of their liking, named Tamao Kikunoi.
Called the "infamous Marie Antoinette of the East", Miss Nakahara is extremely wealthy and usually travels unconventionally, such as in a fleet of helicopters.
The Landlady is rarely around (she is usually in France with her latest boyfriend or fiancé) and the four males have managed to convince her that Sunako is on the fast track to becoming a lady when in reality she is not.
The Landlady is the younger sister of Sunako's father and, like her niece, requires very little effort to frighten the boys.
Voiced by: Mai Nakahara (Japanese); Jessica Boone (English) Live-Action: Ranko Kobe Noi Kasahara (笠原 乃依, Kasahara Noi) is a very pretty and exceedingly popular girl who becomes Sunako's closest female friend at school.
Voiced by: Norihisa Mori (Japanese); Matt Greenfield (English) Hiroshi-Kun is an anatomical figure, Sunako's most precious possession and best friend.
Sunako takes good care of Hiroshi-Kun, along with her other anatomical friends (Josephine, Akira-kun, and Sergeant George).
The boys are not surprised to discover that Sunako's father is physically huge and not particularly attractive and are more worried about his protective streak towards his daughter, who became increasingly distant from him and refused to let him into her room.
However, after she wins an athletic competition for their class, they come to respect and admire her, though they remain somewhat frightened of her more morbid habits.
In chapter 55, she moved to Florida, where it was shown that Yuki got jealous over a photo she took with an American, and vows to become more of a gentleman himself, which unfortunately resulted in a misunderstanding between the two.
Voiced by: Tsuyoshi Koyama (Japanese); John Swasey (English) The self-proclaimed "Head of the National Boss Association", a street gangster from Kumagoroshi who falls for the SD Sunako, much to the surprise of everyone surrounding him.
Because Yuki left the house during their younger ages, he was rarely around them as they were growing up, and is shown to want to be a big brother to them now, despite it often causing misunderstandings with them.