Opening with the main character, Rahm, waking from stasis, the book covers his campaigns as part of the Tzen Warrior caste against a coalition of large Insect species; the Wasps, Leapers, and Ants, in that order.
However, because of Tzen military practices, that a task force leaves after a predicted proportion of units return, assuming that all others are casualties, Rahm's flight group was stranded on the planet.
While stranded, the team made a number of discoveries, chief among which was the fact that the Tzen needed to find a natural predator of the Leapers to defeat them.
The Leapers are immune to this, however, because they are all capable of reproduction, and bury their eggs, making them impossible to eradicate without levying resources unavailable to the Tzen Empire.
In the middle of the battle, Rahm discovers that, while they have eliminated the Ants primary power source, they had a reserve strong enough for one use of a power-consuming device.