Fillmore travels through different locations, riding his horse that needs to be fed carrots from the farms of in-game friends or bought by the player at the village store.

One night, while riding alone through the meadows, Fenimore Fillmore arrives at a farm that is under attack by gunmen and becomes a hero for helping the farmer, a man named Bannister, fend them off.

Early the next morning, Fenimore awakens to discover that his gun is missing; he assumes that Bannister's son Billy has borrowed it to take it to school.

In his quest to retrieve his weapon, Fenimore becomes involved in helping Bannister and the other farmers in their ongoing fight against Starek, a powerful rancher whose niece, Rhiannon, becomes the love of his life.

In France, the game was localized by Focus Home Interactive in an effort to capitalize on its success with Runaway: A Road Adventure.

[5] However, according to Christopher Kellner of German publisher DTP Entertainment, the game's sales had fallen "a bit short of our expectations" in Germany by January 2005.