The Turkmenian kulan has been reintroduced to Kazakhstan and Uzbekistan, as well as Israel, where the subspecies are hybridizing with Persian onagers in the wild.
No other data existed on the condition of the Turkmenistan populations, but hope remained that small groups of animals still resided in inaccessible areas around Badkhyz, and were thriving in the West Kopetdagh (Sumbar-Chandyr Valley) and Ustyurt Plateau around Lake Sarakamish.
The Transcaspian wild ass is characterized by a pale brown coat, a dark stripe down the spine and white patches on the sides, back and belly.
The genus appears to have spread quickly into the Old World, with the similarly aged Equus livenzovensis documented from western Europe and Russia.
[8] Molecular phylogenies indicate the most recent common ancestor of all modern equids (members of the genus Equus) lived ~5.6 (3.9–7.8) mya.
[9] The oldest divergencies are the Asian hemiones (subgenus E. (Asinus), including the kulan, onager, and kiang), followed by the African zebras (subgenera E. (Dolichohippus), and E. (Hippotigris)).
[10] The Turkmenian kulan lives in Central Asian deltas, hot and cold deserts or semi-deserts, steppes, arid grasslands and shrublands.
The Turkmen specimen used to be the most widespread of the onagers, ranging from northernmost Iran and northern Afghanistan, the Transcaspian Oblast and western China to Ukraine and southern Siberia.
The dominant stallion defends the surrounding areas to the water sources and tries to mate with any females that come close to drink.
The scorching heat of the Central Asian deserts makes the Transcaspian wild ass mostly active at dawn and dusk when temperatures are milder.
[citation needed] Like all onagers, the Turkmenian kulan is threatened by poaching, hunting for their meat and their coat, habitat loss and relationships with apex predators such as Persian leopards, striped hyenas and mostly Himalayan wolves.
On 24 October 2017, nine kulans were taken from Altyn-Emel and released to Altyn Dala protected area of the central Kazakh steppes.
[13] A July 2023 Wired story reported that at least two kulan foals had been born at Tarutino Steppe preserve, and despite the disruptions caused by nearby military activity, the herd has adapted well.