Tylosis (botany)

In woody plants, a tylosis (plural: tyloses) is a bladder-like distension of a parenchyma cell into the lumen of adjacent vessels.

The term tylosis summarises the physiological process and the resulting occlusion in the xylem of woody plants as response to injury or as protection from decay in heartwood.

Observed in section under a microscope, tyloses appear as balloon-like protrusions emanating from axial paratracheal parenchyma cells into xylem vessels through pits linking the two.

[3] These blockages can be used in addition to gum plugs as soon as vessels become filled with air bubbles, and they help to form a stronger heartwood by slowing the progress of rot.

The effectiveness of both vertical and horizontal barriers is affected by the speed at which they are established by tyloses, being typically faster in healthier plants.

Section of Carpinus betulus wood showing the distinctive dark areas created as a result of tylosis in response to fungal infection