Velociraptor (commonly referred to as "raptor") is one of the dinosaur genera most familiar to the general public due to its prominent role in the Jurassic Park films.
In reality, however, Velociraptor was roughly the size of a turkey, considerably smaller than the approximately 2 m (6.6 ft) tall and 90 kg (200 lb) reptiles seen in the novels and films (which were based on members of the related genus Deinonychus).
During an American Museum of Natural History expedition to the Flaming Cliffs (Bayn Dzak or Bayanzag) of the Djadochta Formation, Gobi Desert, on 11 August 1923, Peter Kaisen discovered the first Velociraptor fossil known to science—a crushed but complete skull, associated with one of the raptorial second toe claws (AMNH 6515).
[3] While North American teams were shut out of communist Mongolia during the Cold War, expeditions by Soviet and Polish scientists, in collaboration with Mongolian colleagues, recovered several more specimens of Velociraptor.
[13] In 1999, Rinchen Barsbold and Halszka Osmólska reported a juvenile Velociraptor specimen (GIN or IGM 100/2000), represented by a complete skeleton including the skull of a young individual.
[19] In 2021 Powers with team used Principal Component Analysis to separate dromaeosaurid maxillae, most notably finding that MPC-D 100/982 falls outside the instraspecific variability of V. mongoliensis, arguing for a distinct species.
An elongated, near oval-shaped hole was developed in the center of the lower jaw (the mandibular fenestra), and it was produced by the joint of the dentary, surangular, and angular bones.
However, at least one specimen has preserved a series of intact tail vertebrae curved sideways into an S-shape, suggesting that there was considerably more horizontal flexibility than once thought.
[34] Several studies published during the 2010s, including expanded versions of the analyses that found support for Velociraptorinae, have failed to resolve it as a distinct group, but rather have suggested it is a paraphyletic grade which gave rise to the Dromaeosaurinae.
[27] Examinations of the endocranium of Velociraptor indicate that it was able to detect and hear a wide range of sound frequencies (2,368–3,965 Hz) and could track prey with ease as a result.
For instance, most Asian species have elongated snouts based on the maxilla (namely velociraptorines), indicating a selective feeding in Velociraptor and relatives, such as picking up small, fast prey.
[43] The "Fighting Dinosaurs" specimen, found in 1971, preserves a Velociraptor mongoliensis and Protoceratops andrewsi in combat and provides direct evidence of predatory behavior.
[46] In the "Fighting Dinosaurs" specimen, the Velociraptor lies underneath, with one of its sickle claws apparently embedded in the throat of its prey, while the beak of Protoceratops is clamped down upon the right forelimb of its attacker.
This suggests Velociraptor may have used its sickle claw to pierce vital organs of the throat, such as the jugular vein, carotid artery, or trachea (windpipe), rather than slashing the abdomen.
These researchers proposed that, like accipitrids, the dromaeosaur would then begin to feed on the animal while it was still alive, and prey death would eventually result from blood loss and organ failure.
This proposal is based primarily on comparisons between the morphology and proportions of the feet and legs of dromaeosaurs to several groups of extant birds of prey with known predatory behaviors.
Fowler found that the feet and legs of dromaeosaurs most closely resemble those of eagles and hawks, especially in terms of having an enlarged second claw and a similar range of grasping motion.
The arms, which could exert a lot of force but were likely covered in long feathers, may have been used as flapping stabilizers for balance while atop a struggling prey animal, along with the stiff counterbalancing tail.
The authors concluded that the find represented "late-stage carcass consumption by Velociraptor" as the predator would have eaten other parts of a freshly killed Protoceratops before biting in the jaw area.
[37] In 2023, Seishiro Tada and team examined the nasal cavities of ectotherm (cold-blooded) or endotherm (warm-blooded) species, in order to evaluate the thermoregulatory physiology of non-avian dinosaurs compared to these groups.
Tada with team suggested that Velociraptor and most other non-avian dinosaurs may not have possessed a fully or well-developed nasal thermoregulation apparatus as modern endothermic animals do.
[56] Norell with colleagues in 1995 reported one V. mongoliensis skull bearing two parallel rows of small punctures on its frontal bones that, upon closer examination, match the spacing and size of Velociraptor teeth.
[58] In 2012 David Hone and team reported another injured Velociraptor specimen (MPC-D 100/54, roughly a sub-adult individual) found with the bones of an azhdarchid pterosaur within its stomach cavity, was carrying or recovering from an injury sustained to one broken rib.
[59] V. osmolskae lived alongside the ankylosaurid Pinacosaurus mephistocephalus; alvarezsaurid Linhenykus; closely related dromaeosaurid Linheraptor; oviraptorids Machairasaurus and Wulatelong; protoceratopsids Bagaceratops and Protoceratops hellenikorhinus; and troodontids Linhevenator, Papiliovenator, and Philovenator.
[61] The abundant sediments—sands, sandstones, or caliche—of the Djadochta Formation were deposited by eolian (wind) processes in arid settings with fields of sand dunes and only intermittent streams, as indicated by very sparse fluvial (river-deposited) sedimentation, under a semi-arid climate.
[3][61] The younger Turgrugyin Member (mainly Tugriken Shireh locality) has produced the bird Elsornis; dromaeosaurid Mahakala: ornithomimid Aepyornithomimus; and protoceratopsid Protoceratops andrewsi.
The type specimen was discovered at the Flaming Cliffs site (sublocality of the larger Bayn Dzak locality/region),[3] while the "Fighting Dinosaurs" were found at the Tugrik Shire locality (also known as Tugrugeen Shireh and many other spellings).
Based on deposits (such as structureless sandstones), it has been concluded that a large number of specimens were buried alive during powerful sand-bearing events, common to these paleoenvironments.
[66] Velociraptor is commonly perceived as a vicious and cunning killer thanks to their portrayal in the 1990 novel Jurassic Park by Michael Crichton and its 1993 film adaptation, directed by Steven Spielberg.
[67] Crichton met with the discoverer of Deinonychus, John Ostrom, several times at Yale University to discuss details of the animal's possible range of behaviors and appearance.