Cuckmere Valley is a civil parish in the Wealden District of East Sussex, England.

As its name suggests, the parish consists of a number of small settlements in the lower reaches of the River Cuckmere.

The church, dedicated to the Good Shepherd, is believed to be the smallest in England, having been rebuilt using only the original chancel after a fire had destroyed the remainder of the building.

After the Norman conquest, the village was given to Robert, Count of Mortain, half brother of William the Conqueror.

It was built by Benedictine monks in 1280 and continuously occupied since that period although the clergy vacated in the 1970s and it is now a private home.

In addition, there are three Sites of Special Scientific Interest within the parish; Seaford to Beachy Head, Wilmington Downs and Lullington Heath.

[16] Wilmington Downs is another area of biological interest; its chalk grassland is habitat for several rare species.