The eldest son of William Windham of Earsham (died 1730) and Anne Tyrrell, he came of an old Norfolk family.
He is sometimes called William Windham Ashe, a misnomer according to the History of Parliament.
He was elected a Fellow of the Royal Society in 1749 although his fellowship lapsed after only two years.
In April 1734, he had married Mary, Dowager Countess of Deloraine (d. 1744), daughter of Capt.
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