Ypupiara (meaning "the one who lives in the water") is an extinct genus of unenlagiine theropod from the Late Cretaceous Serra da Galga Formation of Brazil.

The holotype, DGM 921-R, a right maxilla and dentary (which was associated with a fish jaw), was discovered in a layer of the Late Cretaceous Serra da Galga Formation of Brazil.

It was found by Alberto Lopa sometime in the 1950s, possibly in 1957, after which Llewellyn Ivor Price listed the fossil as belonging to an indeterminate vertebrate.

[1] The generic name, Ypupiara, is derived from a Tupi word meaning "the one who lives in the water," in reference to a local mythological creature and its inferred diet of fish.

[1] The describers of Ypuparia suggested that unenlagiines such as Ypupiara and its sister taxon Austroraptor likely consumed fish for a considerable part of their diet, based on their non-serrated conical teeth that are similar to those of other piscivorous tetrapods including gavialoid crocodylians, spinosaurid theropods, and anhanguerid pterosaurs.