It saw active service in France and on the Eastern Front and was destroyed in fighting in Romania in August 1944.

The 335th Infantry Division was formed in Konstanz in November 1940 under the command of Generalleutnant[Note 1] Max Dennerlein.

Following the Allied invasion of French North Africa in November 1942, German forces moved into Vichy France in an operation known as Case Anton.

Transferred to Romania later that year,[1] it received some personnel from the recently disbanded 5th Field Division (L).

The division, commanded by Oberst Eugen Franz Brechtel, was destroyed and its survivors surrendered to the Soviets.