The holotype fossil was recovered from Resner Canyon, location National Museum of Natural History USNM 11018 in the Toroda Creek Graben of Northeast Washington State.

[1] The paratype specimen was recovered from sediments of the Ypresian Allenby Formation exposed in the Blue Flame mine near Princeton, British Columbia.

Both the Allenby and Klondike Mountain Formations preserve upland temperate floras which were first interpreted as being Microthermal,[1] however further study has shown them to be more mesothermal in nature.

[2] The plant community preserved in the Klondike Mountain formation is a mixed confer-broad leaf forest with large pollen elements of birch and golden larch, but also having notable traces of fir, spruce, cypress, and palm.

[1] The specimens were studied by paleobotanists Jack A. Wolfe of the United States Geological Survey, Denver office and Toshimasa Tanai of Hokkaido University.