One of the two specimens was recovered from sediments of the early Eocene, Ypresian[2] Allenby Formation, exposed at the One Mile Creek site near Princeton, British Columbia.

The other fossil was recovered from the "Corner lot site", Burke Museum of Natural History and Culture location UWBM A0307 in Republic, Washington.

Location UWBM A0307 is an outcropping of shale belonging to the Ypresian Klondike Mountain Formation and is in the Republic Graben of Northeast Washington State.

[2] The plant community preserved in the Klondike Mountain formation is a mixed confer-broad leaf forest with large pollen elements of birch and golden larch, but also having notable traces of fir, spruce, cypress, and palm.

[1] The specimens were studied by paleobotanists Jack A. Wolfe of the United States Geological Survey, Denver office and Toshimasa Tanai of Hokkaido University.