Ambisporaceae Archaeosporaceae Geosiphonaceae Archaeosporales is an order of fungi best known as arbuscular mycorrhiza to vascular land plants (Tracheophyta).

But also form free living endocyte symbioses with cyanobacteria.

[1] The free living forms may have a Precambrian fossil record back 2.2 Ga, well before evolution of Tracheophyta.

[4] However, the earliest fossils of Opisthokonta otherwise date back to the early Tonian, thus making this possibility questionable.

Data related to Archaeosporales at Wikispecies