Aulacostephanus is an extinct ammonoid cephalopod genus from the Upper Jurassic Tithonian belonging to the perisphinctoidean family Aulacostephanidae.

Aulacostephanus produced a discoidal, strongly ribbed, evolute shell of moderate size, reaching diameters of 16 cm (~6.5in.

Sides are flat, the venter steeply rounded, with an open bend or median groove running down the middle.

Clutches of eggs attributed to this genus have been discovered in the Kimmeridge Clay.

[2] Jurassic sediments of Germany, the Russian Federation, Switzerland and the United Kingdom [3]