There is a correlated relationship between how territorial an individual is, how likely they are to explore their environment, and what strategy they use to avoid predation.
Through the 90s, it was infrequently used analogously for "personality" in some behavioral ecology literature, but it was still primarily used in psychology literature to describe inter-individual differences in behavior to model systems like rodents and primates.
Often, behavior is considered infinitely plastic and can easily be adapted to changing environmental conditions.
However, the phenotypic correlations that form behavioral types do not necessarily signify that they have a genetic basis.
Without a genetic basis, behavioral syndromes must be the result of environmental conditions.
[11] The simplest way for a behavioral syndrome to form is through a genetic polymorphism, meaning two or more alleles at the same locus.
In one of the best documented examples of this, a single gene (for) controls the foraging distance and a suit of related traits in Drosophila melanogaster.
[14] Non-genetic behavioral syndromes have received almost no focus in recent years, although studies of associated traits that appear to be environmentally determined are not uncommon under other names.