Columella (botany)

Columella (in plants) is an axis of sterile tissue which passes through the center of the spore-case of mosses.

[1] In fungi, it refers to a centrally vacuolated part of a hypha, bearing spores.

The word finds analogous usage in myxomycetes.

[citation needed] The term columella is also used to refer to story 1 to story 4 (S1 – S4) cells in the root cap, located apically of the quiescent centre.

[citation needed] In vascular plants, columella tissue helps the roots to perceive gravity.