It is the sister taxon of Discomedusae and contains about 50 named species, all included in the order Coronatae.

[2] The mouthparts of the members of this subclass are characterised by simple lips on a short manubrium as well as by robust marginal tentacles that arc away from the mouth.

It is likely that these marginal tentacles are used to capture plankton, including crustaceans and small fish in the larger species.

[3] Coronatae is the only order in this subclass, with around thirty-five species having been described, most of them deepwater jellyfish.

The mouth opens into a central stomach which is divided into four pockets by septa, on which are located the gastric filaments and the eight gonads.